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Two years later, Dal Sasso et al. More recently, the enamel texture of Spinosaurus teeth from the Kem Kem was investigated by Hasegawa et al. Spinosaurus aegyptiacus was revealed to be a semi-aquatic quadrupedal theropod with short hind limbs and dense postcranial bones . Based on the newly discovered material of Spinosaurus , Ibrahim et al. The recent description of additional material of the spinosaurid Sigilmassasaurus brevicollis from the Kem Kem beds by Evers et al. The functional morphology of the spinosaurid skull was investigated by Therrien et al. The capture of small prey items by Suchomimus was possible thanks to its upturned chin with the terminal rosette, large mandibular symphysis allowing to resist the important stresses induced by struggling prey, and conical teeth designed to impale and hold prey and withstand bending loads applied in all directions . Using FEA on 2D models of skulls, Rayfield  found that Suchomimus and Spinosaurus skulls experience cranial stresses in different ways. Whereas Suchomimus is scaling in a similar manner to most non-spinosaurid tetanurans, Spinosaurus experiences a much higher magnitude of cranial stress than what would be predicted, suggesting it may have fed on smaller prey. Here we report additional cranial material of spinosaurids consisting of six isolated quadrates most likely coming from the Kem Kem beds. The quadrate is a cranial bone of endochondral origin that articulates with the mandible in all gnathostomes other than mammals [50’52]. In theropods, the quadrate had many important functions such as a structural support for the basicranium, an articulatory element with the lower jaw, an insertion area for several muscles, and in hosting important nerves, pneumatic sinuses, and vascular passages (e. Material and geological settings. Six isolated quadrates of different sizes were collected by locals and acquired commercially in the Kem Kem region of southeastern Morocco (Fig 1). KK378; Figs 2’4; Fig C in S1 File) were provided by Francois Escuillie and are deposited in the collections of the Museum d’Histoire Naturelle of Marrakech (MHNM; see S1 File for a list of institutional abbreviations), casinos in pretoria.
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